If you think you might need to defragment

  1. Try restarting first. It might help, and it’s easy to do.
  2. If your disks are almost full, and you often modify or create large files (such as editing video, but see the Tip below if you use iMovie and Mac OS X 10.3), there’s a chance the disks could be fragmented. In this case, you might benefit from defragmentation, which can be performed with some third-party disk utilities.
  3. Another option is to back up your important files, erase the hard disk, then reinstall Mac OS X and your backed up files.

Reference: About disk optimization with Mac OS X.

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